Dr Frank Glaw and Professor Miguel Vences wrote the first edition of the definitive guide to Madagascar\u2019s Amphibians and Reptiles in 1992. Since then, we have had two new versions (1994, 2007). The latest of these is nearly a decade out of date now, and contains many undescribed taxa. While it is not within my ability to direct you in the adding of all of the newly described species to your copy of this book, I here list the species that were not yet described but were figured<\/strong><\/em> in that book (listed by page number, and species number on that page). There are dozens of species completely missing from it, and these are not mentioned here.<\/p>\n Note: I maintain this page periodically. It is neither 100% complete, nor am I liable for fallacies it might contain. It also does not contain species described since 2007 that were not figured in the guide. For a complete list of species known from Madagascar, click here<\/a>.<\/p>\n This page was last updated\u00a0 p. 108, sp. 3: Scaphiophryne matsoko<\/em> Raselimanana, Raxworthy, Andreone, Glaw & Vences, 2014<\/p>\n p. 110, sp. 1: Scaphiophryne brevis<\/em> was described in Boulenger 1896 as Calophrynus brevis<\/em>, not by Grandidier.<\/p>\n p. 110, sp. 2:\u00a0Scaphiophryne obscura<\/em> (Grandidier, 1872)<\/p>\n p. 110, sp. 3: Scaphiophryne calcarata<\/em> (Mocquard, 1895)<\/p>\n p. 118, sp. 1:\u00a0Rhombophryne testudo<\/em>: records from Andapa and Sambava refer to\u00a0R. coudreaui<\/em> or an undescribed related species.<\/p>\n p. 118, sp. 2: Rhombophryne coudreaui<\/em>: correct attribution (Angel, 19<\/strong>38), not 1838.<\/p>\n p. 118, sp. 3:\u00a0Rhombophryne serratopalpebrosa<\/em>: known only from the holotype. Records of ‘other specimens from Marojejy (fig. 3a)’ refer to\u00a0R. vaventy\u00a0<\/em>Scherz, Ruthensteiner, Vences & Glaw, 2014. ‘Specimens from Anjanaharibe (fig. 3b)’ refer to\u00a0R. regalis<\/em>.<\/p>\n p. 118, sp. 5: Rhombophryne guetherpetersi<\/em>: morphology unclear but under review by us. Tibiotarsal articulation reaches the insertion of the arms; possessing large tibial and dorsolateral glands; weak superciliary spines!<\/p>\n p. 118, fig. \u201c5?\u201d above: Rhombophryne ornata<\/em> Scherz, Ruthensteiner, Vieites, Vences & Glaw, 2015<\/p>\n p. 118, fig. \u201c5?\u201d below: probably\u00a0Rhombophryne tany<\/em> Scherz, Ruthensteiner, Vieites, Vences & Glaw, 2015<\/p>\n p. 119, fig. 3a:\u00a0Rhombophryne vaventy<\/em> Scherz, Ruthensteiner, Vences & Glaw, 2014<\/p>\n p. 119, fig. 3b: Rhombophryne regalis<\/i> Scherz,\u00a0Hawlitschek, Andreone, Rakotoarison, Vences & Glaw, 2017<\/p>\n p. 119, fig. 5: not Rhombophryne guentherpetersi<\/em>. Probably a new species.<\/p>\n p. 120, sp. 1: undescribed\u00a0Rhombophryne\u00a0<\/em>species (also figured on p. 121 1a and 1b); name ‘alluaudi’<\/em> transferred to genus\u00a0Plethodontohyla<\/em> by Bellati et al. 2018\u00a0[pdf<\/a>]. Map pertaining to this species is useless, because it does not distinguish\u00a0Rhombophryne<\/em> and\u00a0Plethodontohyla<\/em> lineages.\u00a0Plethodontohyla alluaudi<\/em> is restricted to southeastern Madagascar, from Ranomafana region down to Tolagnaro.<\/p>\n p. 120, sp. 2: Rhombophryne mangabensis<\/em> Glaw, K\u00f6hler & Vences, 2010<\/p>\n p. 124, sp. 6: Plethodontohyla fonetana<\/em> Glaw, K\u00f6hler, Bora, Rabibisoa, Ramilijaona & Vences, 2007<\/p>\n p. 126, sp. 1:\u00a0Stumpffia tetradactyla<\/em> from Nosy Mangabe refers to\u00a0S. analanjirofo<\/em>\u00a0Rakotoarison,\u00a0Scherz,\u00a0Glaw, K\u00f6hler, Andreone, Franzen, Glos, Hawlitschek, Jono, Mori, Ndriantsoa, Rasoamampionona, Riemann, R\u00f6del, Rosa, Vieites, Crottini & Vences, 2017\u00a0[highres pdf (72 MB)<\/a>|lowres pdf (15 MB)<\/a>]<\/p>\n p. 126, sp. 2:\u00a0Stumpffia tridactyla<\/em> from Andasibe refers to\u00a0S. obscoena\u00a0<\/em>Rakotoarison,\u00a0Scherz,\u00a0Glaw, K\u00f6hler, Andreone, Franzen, Glos, Hawlitschek, Jono, Mori, Ndriantsoa, Rasoamampionona, Riemann, R\u00f6del, Rosa, Vieites, Crottini & Vences, 2017 [highres pdf (72 MB)<\/a>|lowres pdf (15 MB)<\/a>]<\/p>\n p. 126, spp. 3,4: moved to Anilany<\/em> Scherz, Vences, Rakotoarison, Andreone, K\u00f6hler, Glaw & Crottini, 2016 by Scherz et al. 2016 Mol. Phyl. Evol. 100:372<\/p>\n p. 128, sp. 3:\u00a0Stumpffia gimmeli<\/em> refers to multiple different species; see Rakotoarison et al. 2017\u00a0[highres pdf (72 MB)<\/a>|lowres pdf (15 MB)<\/a>]<\/p>\n p. 128, sp. 4: refers to the genus Mini<\/em> Scherz, Hutter, Rakotoarison, Riemann, R\u00f6del, Ndriantsoa, Glos, Roberts, Crottini, Vences & Glaw, 2019. The record refers to two of the three species of this genus:\u00a0Mini mum<\/em> Scherz et al., 2019 from Manombo, and\u00a0Mini scule<\/em> Scherz et al., 2019 from Sainte Luce. A third species,\u00a0Mini ature<\/em> Scherz et al., 2019 is also present in Andohahela. See Scherz et al. 2019 [PDF<\/a>].<\/p>\n p. 130, sp. 5: Stumpffia kibomena<\/em> Glaw, Vallan, Andreone, Edmonds, Dolch & Vences, 2015<\/p>\n p. 130, sp. 6:\u00a0Stumpffia nigrorubra<\/em>\u00a0Rakotoarison,\u00a0Scherz,\u00a0Glaw, K\u00f6hler, Andreone, Franzen, Glos, Hawlitschek, Jono, Mori, Ndriantsoa, Rasoamampionona, Riemann, R\u00f6del, Rosa, Vieites, Crottini & Vences, 2017 [highres pdf (72 MB)<\/a>|lowres pdf (15 MB)<\/a>]<\/p>\n p. 130, sp. 7:\u00a0Stumpffia diutissima<\/em>\u00a0Rakotoarison,\u00a0Scherz,\u00a0Glaw, K\u00f6hler, Andreone, Franzen, Glos, Hawlitschek, Jono, Mori, Ndriantsoa, Rasoamampionona, Riemann, R\u00f6del, Rosa, Vieites, Crottini & Vences, 2017 [highres pdf (72 MB)<\/a>|lowres pdf (15 MB)<\/a>]<\/p>\n p. 130, sp. 8:\u00a0Stumpffia achillei<\/em>\u00a0Rakotoarison,\u00a0Scherz,\u00a0Glaw, K\u00f6hler, Andreone, Franzen, Glos, Hawlitschek, Jono, Mori, Ndriantsoa, Rasoamampionona, Riemann, R\u00f6del, Rosa, Vieites, Crottini & Vences, 2017 [highres pdf (72 MB)<\/a>|lowres pdf (15 MB)<\/a>]<\/p>\n p. 135, fig. 5d: Cophyla puellarum<\/em> Rakotoarison, Crottini, M\u00fcller, R\u00f6del, Glaw & Vences, 2015. 27.3\u201333.6 mm SVL, yellowish green venter, toes 1<2<3<5<4, fingers 1<2=4<3, tibio-tarsal articulation between forelimb and tympanum, call 326\u2013390 ms 3\u20134 times per 10 seconds.<\/p>\n p. 136, sp. 2:\u00a0Platypelis ranjomena <\/em>Glaw, Scherz, Rakotoarison, Crottini, Raselimanana, Andreone, K\u00f6hler & Vences, 2020<\/p>\n p. 136, sp. 5: Platypelis ravus<\/em> Glaw, K\u00f6hler & Vences, 2012, 17\u201319 mm SVL<\/p>\n p. 138, sp. 3: records from Marojejy refer to\u00a0Cophyla fortuna\u00a0<\/em>Rakotoarison, Scherz, Bletz, Razafindraibe, Glaw & Vences, 2019. Other records from the north east probably refer to additional undescribed species.<\/p>\n p. 140, sp. 4: Anodonthyla emilei<\/em> Vences, Glaw, K\u00f6hler & Wollenberg, 2010<\/p>\n p. 142, sp. 3: Anodonthyla vallani <\/em>Vences, Glaw, K\u00f6hler & Wollenberg, 2010<\/p>\n p. 142, sp. 4: Anodonthyla<\/em> jeanbai<\/em> Vences, Glaw, K\u00f6hler & Wollenberg, 2010<\/p>\n p. 148, sp. 5: Boophis calcaratus<\/em> Vallan, Vences & Glaw, 2010<\/p>\n p. 152, sp. 3: Boophis androngoloaka<\/em> (Ahl, 1928) – resurrected by Glaw et al. 2010 Zootaxa 2383:1-82<\/p>\n p. 152, sp. 4: Boophis piperatus<\/em> Glaw, K\u00f6hler, de la Riva, Vieites & Vences, 2010<\/p>\n p. 154, sp. 4: Boophis narinsi<\/em> Vences, Gehara, K\u00f6hler & Glaw, 2012<\/p>\n p. 154, sp. 5: Boophis arcanus<\/em> Glaw, K\u00f6hler, de la Riva, Vieites & Vences, 2010<\/p>\n p. 160, sp. 2: Boophis quasiboehmei<\/em> Vences, K\u00f6hler, Crottini & Glaw, 2010<\/p>\n p. 160, sp. 4: Boophis popi<\/em> K\u00f6hler, Glaw, Rosa, Gehring, Pabijan, Andreone & Vences, 2011<\/p>\n p. 162, sp. 3: Boophis brachychir<\/em> (Boettger, 1882) – resurrected by Glaw et al. 2010 Zootaxa 2383:1-82<\/p>\n p. 162, sp. 4: Boophis entingae<\/em> Glaw, K\u00f6hler, de la Riva, Vieites & Vences, 2010<\/p>\n p. 164, sp. 1: Boophis roseipalmatus<\/em> Glaw, K\u00f6hler, de la Riva, Vieites & Vences, 2010<\/p>\n p. 164, sp. 2: Boophis spinophis<\/em> Glaw, K\u00f6hler, de la Riva, Vieites & Vences, 2010<\/p>\n p. 164, sp. 4: Boophis obscurus<\/em> (Boettger 1913) – resurrected by Glaw et al. 2010 Zootaxa 2383:1-82<\/p>\n p. 166, sp. 1: Boophis tsilomaro<\/em> Vences, Andreone, Glos & Glaw, 2010<\/p>\n p. 166, sp. 4: Boophis praedictus<\/em> Glaw, K\u00f6hler, de la Riva, Vieites & Vences, 2010. Known from Makira, Masoala (see next comment) and Vevembe;\u00a0Vences et al. 2010 Zootaxa 2544:54.<\/p>\n p. 166, sp. 5: According to Vences et al. 2010 Zootaxa 2544:54, this ‘species’ probably just represents juveniles of\u00a0B. praedictus<\/em>, as the depicted individual on page 167 is identical in 16S rRNA sequence to a specimen of\u00a0B. praedictus<\/em> from Makira.<\/p>\n p. 168, sp. 4: Boophis sandrae<\/em> Glaw, K\u00f6hler, de la Riva, Vieites & Vences, 2010<\/p>\n p. 170, sp. 2:\u00a0Boophis tampoka<\/em> K\u00f6hler, Glaw & Vences, 2007\u2014additional localities in northern Madagascar that are somewhat strongly genetically differentiated (Antambato, Antsohihy-Befandriana, Sahaovy, and Antsatramidola) were reported in\u00a0Vences et al. 2011 Herpetology Notes 4:239.<\/p>\n p. 172, sp. 3: Boophis boppa<\/em> Hutter, Lambert, Cobb, Andrianpanonanana & Vences, 2015<\/p>\n p. 172, sp. 4: Boophis miadana <\/em>Glaw, K\u00f6hler, de la Riva, Vieites & Vences, 2010<\/p>\n p. 172, sp. 5: Boophis haingana <\/em>Glaw, K\u00f6hler, de la Riva, Vieites & Vences, 2010<\/p>\n p. 172, sp. 8: Boophis luciae <\/em>Glaw, K\u00f6hler, de la Riva, Vieites & Vences, 2010<\/p>\n p. 174, spp. 1 & 2 were moved to the Boophis ulftunni<\/em> species group, together with B. baetkei<\/em> K\u00f6hler, Glaw & Vences, 2008 (top left figure on p. 446), but were then placed into the\u00a0B. majori<\/em> species group by Hutter et al. 2018<\/a>.<\/p>\n p. 174, sp. 1: Boophis ulftunni<\/em> Wollenberg, Andreone, Glaw & Vences, 2008<\/p>\n p. 174, sp. 2: Boophis lilanae<\/em> K\u00f6hler, Glaw & Vences, 2008<\/p>\n p. 182, sp. 2: Aglyptodactylus australis<\/em> K\u00f6hler, Glaw, Pabijan & Vences, 2015<\/p>\n p. 182, sp. 3: Aglyptodactylus inguinalis<\/em> (G\u00fcnther, 1877) – resurrected by K\u00f6hler et al. 2015 Zootaxa 4006:401<\/p>\n p. 182, sp. 4: Aglyptodactylus chorus<\/em> K\u00f6hler, Glaw, Pabijan & Vences, 2015<\/p>\n p. 186, sp. 3: Blommersia dejongi<\/em> Vences, K\u00f6hler, Pabijan & Glaw, 2010<\/p>\n p. 186, sp. 4: Blommersia variabilis<\/em> Pabijan, Gehring, K\u00f6hler, Glaw & Vences, 2011<\/p>\n p. 186, sp. 5: Blommersia galani<\/em> Vences, K\u00f6hler, Pabijan & Glaw, 2010<\/p>\n p. 202, sp. 5:\u00a0Guibemantis wattersoni<\/em> Lehtinen, Glaw & Vences, 2011<\/p>\n p. 204, sp. 5: Guibemantis tasifotsy<\/em> Lehtinen, Glaw, Andreone, Pabijan & Vences, 2012<\/p>\n p. 204, sp. 6: Records of\u00a0G. liber<\/em> from the northeast refer to\u00a0Guibemantis razandry <\/em>Koppetsch, Pabijan, Hutter, K\u00f6hler, Gehring, Rakotoarison, Ratsoavina,\u00a0Scherz<\/strong>, Vieites, Glaw & Vences, 2023,\u00a0G. razoky<\/em> Koppetsch et al. 2023, and\u00a0G. fotsitenda<\/em> Koppetsch et al. 2023. Only\u00a0G. razoky<\/em> is present on Montagne d’Ambre, but most other sites have two species in sympatry. See the Koppetsch et al. paper here<\/a>.<\/p>\n p. 206, sp. 2: records from Marojejy may refer to Spinomantis tavaratra<\/em> Cramer, Rabibisoa & Raxworthy, 2008, and thus photo 2b on p. 207 may be that species.<\/p>\n p. 206, sp. 3:\u00a0Spinomantis<\/em>\u00a0massi<\/strong><\/em>, not massorum<\/em>.<\/p>\n p. 208, sp. 4: photo 4b depicts\u00a0Spinomantis beckei<\/em> Vences, K\u00f6hler & Glaw, 2017.\u00a0S. bertini<\/em> is found only at lower elevations in Andohahela.<\/p>\n p. 210, sp. 1: populations formerly assigned to G. boulengeri<\/em> from Ambatolahy (Fig. 1b and ventral view) are now assigned to G. sergei<\/em> Miralles, K\u00f6hler, Glaw, Wollenberg Valero, Crottini, Rosa, du Preez, Gehring, Vieites, Ratsoavina & Vences, 2023<\/p>\n p. 210, sp. 3:\u00a0Gephyromantis cornucopia<\/em> Miralles, K\u00f6hler, Glaw, Wollenberg Valero, Crottini, Rosa, du Preez, Gehring, Vieites, Ratsoavina & Vences, 2023<\/p>\n p. 216, sp. 1:\u00a0Gephyromantis malagasius<\/em> (Methuen & Hewitt, 1913) due to synonymisation by Vences et al. 2022 [pdf<\/a>]<\/p>\n p. 216, sp. 2: Populations from Ambatovaky, Mananara, and Tampolo are probably\u00a0Gephyromantis matsilo <\/em>Vences, K\u00f6hler, Crottini, Hofreiter, Hutter, du Preez, Preick, Rakotoarison, Rancilhac, Raselimanana, Rosa, Scherz & Glaw, 2022;\u00a0population from Masoala (Fig. 2 on p. 217) is now Gephyromantis oelkrugi <\/em>Vences, K\u00f6hler, Crottini, Hofreiter, Hutter, du Preez, Preick, Rakotoarison, Rancilhac, Raselimanana, Rosa, Scherz & Glaw, 2022. Folohy is type locality of\u00a0G. malagasius<\/em>, and not among localities of these other species.<\/p>\n p. 216, sp. 3:\u00a0Gephyromantis fiharimpe <\/em>Vences, K\u00f6hler, Crottini, Hofreiter, Hutter, du Preez, Preick, Rakotoarison, Rancilhac, Raselimanana, Rosa,\u00a0Scherz\u00a0& Glaw, 2022<\/p>\n p. 216, sp. 6: Gephyromantis ranjomavo<\/em> Glaw & Vences, 2011<\/p>\n p. 218, sp. 2: Gephyromantis atsingy<\/em> Crottini, Glaw, Casiraghi, Jenkins, Mercurio, Randrianantoandro, Randrianirina & Andreone, 2011<\/p>\n p. 218, sp. 4:\u00a0Gephyromantis kintana<\/em> Cocca, Andreone, Belluardo, Rosa, Randrianirina, Glaw & Crottini, 2020 \u2014\u00a0G. azzurrae<\/em> was synonymised with G. corvus<\/em> by these authors.<\/p>\n p. 220, sp. 4: Gephyromantis tahotra<\/em> Glaw, K\u00f6hler & Vences, 2011<\/p>\n p. 222, sp. 2: This account mostly refers to specimens now genetically assigned to\u00a0G. luteus<\/em>.\u00a0G. sculpturatus<\/em> (Ahl, 1929) is now a junior synonym of G. plicifer<\/em>.\u00a0See Vences et al. 2021 [pdf<\/a>]<\/p>\n p. 222, sp. 3: This account mostly refers to specimens belonging to\u00a0G. pedronoi<\/em> Vences, K\u00f6hler, Andreone, Craul, Crottini, du Preez, Preick, Rancilhac, R\u00f6del, Scherz, Streicher, Hofreiter & Glaw, 2021. Real\u00a0G. plicifer<\/em> is only known from Antoetra and Tsinjoarivo. See Vences et al. 2021 [pdf<\/a>]<\/p>\n p. 230, sp. 1: localities are wrong. See Rancilhac et al. 2020<\/a> for correct map of this species. It is only found in the east of Madagascar.<\/p>\n2023-04-30<\/code><\/p>\n